Portal:Amphibians

The Amphibian Portal

Anaschisma

Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods, but excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs and toads), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.

Young amphibians generally undergo metamorphosis from an aquatic larval form with gills to an air-breathing adult form with lungs. Amphibians use their skin as a secondary respiratory interface, and some small terrestrial salamanders and frogs even lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require access to water bodies to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators to habitat conditions; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.

The earliest amphibians evolved in the Devonian period from tetrapodomorph sarcopterygians (lobe-finned fish with articulated limb-like fins) that evolved primitive lungs, which were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (predecessors of mammals). The origin of modern lissamphibians, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. The most popular hypothesis is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. Another hypothesis is that they emerged from lepospondyls. A fourth group of lissamphibians, the Albanerpetontidae, became extinct around 2 million years ago. (Full article...)

Selected frog article

Pipa pipa, the common Suriname toad (Pipidae)

Pipoidea are a clade of frogs, that contains the most recent common ancestor of living Pipidae and Rhinophrynidae as well as all its descendants. It is broadly equivalent to Xenoanura. (Full article...)

List of selected frog articles

Selected salamander article

Alpine salamander (Salamandra atra)

The Salamandroidea are a suborder of salamanders, referred to as advanced salamanders. The members of the suborder are found worldwide except for Antarctica, sub-Saharan Africa, and Oceania. They differ from suborder Cryptobranchoidea as the angular and prearticular bones in their lower jaws are fused, their trunk ribs are bicapitate, and all members use internal fertilization. The female is fertilized by means of a spermatophore, a sperm-containing cap placed by the male in her cloaca. The sperm is stored in spermathecae on the roof of the cloaca until it is needed at the time of oviposition.

The earliest known salamandroid fossils remain contested. Some studies suggest that the earliest salamandroids are represented by specimens of the species Beiyanerpeton jianpingensis and Qinglongtriton gangouensis from the Tiaojishan Formation of Inner Mongolia, China, dated to the Late Jurassic about 157 million years ago. Alternative analyses suggest that Beiyanerpeton jianpingensis and Qinglongtriton gangouensis are stem salamanders and that the oldest known certain salamandroid is Valdotriton gracilis from the Early Cretaceous of Spain, about 127 Ma. (Full article...)

List of selected salamander articles
Caudata Ambystomatidae Cryptobranchoidea Greater siren Salamanders in folklore Sexual selection in amphibians Sirenidae Urodela

Did you know? –

... that the Central American frog Duellmanohyla soralia is critically endangered and its population decline is due, at least in part, to the fungal disease chytridiomycosis?
... that the desert rain frog leaves distinctive footprints on the dunes and a little pile of sand showing the location of its burrow?
... that Pseudophilautus hypomelas, a small shrub frog not seen for more than 130 years and believed to be extinct, was rediscovered in the Peak Wilderness of Sri Lanka?

Selected amphibian type

Spotted salamander, Ambystoma maculatum

Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela, the sole surviving order from the group Caudata. Urodela is a scientific Latin term based on the Ancient Greek οὐρά δήλη: ourà dēlē "conspicuous tail". Caudata is the Latin for "tailed ones", from cauda: "tail".

Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders never have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults.

This group of amphibians is capable of regenerating lost limbs as well as other damaged parts of their bodies. Researchers hope to reverse engineer the regenerative processes for potential human medical applications, such as brain and spinal cord injury treatment or preventing harmful scarring during heart surgery recovery. The remarkable ability of salamanders to regenerate is not just limited to limbs but extends to vital organs such as the heart, jaw, and parts of the spinal cord, showing their uniqueness compared to different types of vertebrates. ⁤⁤This ability is most remarkable for occurring without any type of scarring. ⁤⁤This has made salamanders an invaluable model organism in scientific research aimed at understanding and achieving regenerative processes for medical advancements in human and animal biology. (Full article...)

List of selected amphibian type articles
Frog Toad Caecilian

Selected images

Image 1
Phantasmal poison frog
Photograph: H. Krisp
The phantasmal poison frog (Epipedobates tricolor) is a species of poison dart frog endemic to Ecuador. First described by George Albert Boulenger in 1899, this endangered species is found only in rainforests on the slopes of the Andes.
Image 2
Mantella baroni
Photograph credit: Charles J. Sharp
Mantella baroni, commonly known as Baron's mantella, variegated golden frog or Madagascar poison frog, is a species of frog in the family Mantellidae. It is native to Madagascar and is seen here in Ranomafana National Park in the southeastern part of the country. The species was described in 1888 by George Albert Boulenger, who named it after its collector, Richard Baron. It has been classified as a least-concern species by the International Union for Conservation of Nature due to its relatively wide distribution, but is threatened by habitat loss. The frog's diet includes mites, which cause it to accumulate a high alkaloid concentration in its skin, making it toxic. The bright colours may serve as a warning sign to potential predators of the frog's toxicity.
Image 3
New England Tree Frog
Photo credit: LiquidGhoul
The New England Tree Frog (Litoria subglandulosa) is a species of frog native to the streams of the New England Tablelands, New South Wales, Australia.
Image 4
Dendropsophus microcephalus
Photo: Brian Gratwicke
A male Dendropsophus microcephalus frog, displaying its vocal sac—membranes of skin under the throat or on the corner of the mouth that distend to amplify its call. The species lives in moist areas in most of Central America and northern South America.
Image 5
Dendrobates azureus
Photo: Quartl
Dendrobates azureus, commonly known as the blue poison dart frog or okopipi, is a poison dart frog found in the forests surrounded by the Sipaliwini Savanna in southern Suriname. Authorities have recently treated it as a variant of Dendrobates tinctorius rather than a distinct species as before.
Image 6
Fire salamander
Photo: Archaeodontosaurus
The fire salamander (Salamandra salamandra) is a European salamander species with a long lifespan. These nocturnal animals generally eat various insects, spiders, earthworms and slugs, but they also occasionally eat newts and young frogs.
Image 7
Common toads in amplexus
Photo: Bernie Kohl
Two common toads (Bufo bufo) in amplexus, a form of pseudocopulation found in amphibians and limulids. A male common toad grasps a female with his front legs as part of the mating process. Common toads stay in amplexus for several days. As the female lays a long, double string of small black eggs, the male fertilises them with his sperm; the gelatinous egg strings, which may contain 3000 to 6000 eggs and be 3 to 4.5 metres (10 to 15 ft) in length, are later tangled in plant stalks.
Image 8
Atelopus certus
Photo: Brian Gratwicke
Atelopus certus is a species of toad endemic to the Darién Province of eastern Panama. Much of its range falls within the Darién National Park, a protected area and World Heritage Site. It is classified as endangered, primarily due to the advancing wave of chytridiomycosis moving through Central America.
Image 9
Raorchestes parvulus
Photograph credit: Rushenb
Raorchestes parvulus is a small bush frog native to tropical southeastern Asia. Although the reproductive strategy of this species has not been studied, members of the family typically make a foam nest while mating in vegetation overhanging a stream. They create the foam by beating their legs, and the eggs are deposited into the nest and covered with seminal fluid before the foam hardens into a protective casing. When the eggs hatch, the young push their way out of the nest and fall into the water below.

This picture shows a male R. parvulus frog, photographed in Phu Kradueng National Park, Thailand, calling by inflating its vocal sac.
Image 10
Common Indian Toad
Photo: Lokionly
The Common Indian Toad (Duttaphrynus melanostictus) is native to South Asia. It grows to about 20 cm (8 in) long and breeds during monsoon season. These toads are often seen at night under street lamps especially during times when winged termites swarm. They have been noted to feed on a wide range of invertebrates, including scorpions.
Image 11
Southern brown tree frog
Photograph: JJ Harrison
The southern brown tree frog (Litoria ewingii) is a species of tree frog native to Australia found in a wide range of habitats. Reaching 45 millimetres (1.8 in), this species is generally brown, but green and green-striped morphs are also recorded.
Image 12
Ecnomiohyla rabborum
Photograph: Brian Gratwicke
Rabbs' fringe-limbed treefrog (Ecnomiohyla rabborum) is a large species of frog originally found in the forest canopies of central Panama. Only discovered in 2005, the species is thought to be extinct in the wild; only one specimen, a male at the Atlanta Botanical Garden, survives.
Image 13
Blue Mountains Tree Frog
Photo: Benjamint444
The Blue Mountains Tree Frog (Litoria citropa) is a moderate-sized species of tree frog, up to about 60 mm (2.4 in) in length. It is native to coastal and highland areas of eastern Australia, especially in the Blue Mountains, hence its name.
Image 14
Marsh frog
Photograph credit: Charles James Sharp
The marsh frog (Pelophylax ridibundus) is a species of amphibian native to Europe and parts of Asia. It is the largest type of frog in most of its range, growing to a snout-to-vent length of around 100 mm (4 in); tadpoles can reach up to 190 mm (7.5 in) in length, but this usually occurs in places with long winters where the tadpole has time to grow. Marsh frogs hibernate during the winter, either underwater or in burrows, and are able to use the Earth's magnetic field to locate breeding ponds. This marsh frog was photographed in Kampinos National Park, Poland.
Image 15
Frogspawn
Photo credit: Tarquin
In the life cycle of a frog, a female lays her eggs in a shallow pond or creek, where they will be sheltered from the current and from predators. The eggs, known as frogspawn hatch into tadpoles. The tadpoles develop gradually into adolescent froglets and finally the froglet develops into an adult frog.

Selected toad article

Kihansi spray toad, Nectophrynoides asperginis

Nectophrynoides is a genus of true toads, family Bufonidae. They are endemic to Eastern Arc forests and wetlands in Tanzania, and all except N. tornieri are threatened. Species of the genus are ovoviviparous: fertilization is internal, and the females give birth to fully developed, small toadlets. Together with the West Africa Nimbaphrynoides (which was included in Nectophrynoides in the past) and Limnonectes larvaepartus, they are the only frogs/toads in the world that do not lay eggs. The Ethiopian Altiphrynoides (includes Spinophrynoides), which lay eggs, were also part of Nectophrynoides in the past. (Full article...)

List of selected toad articles

Selected caecilian article

Nectocaecilia petersii is a species of semiaquatic amphibian in the family Typhlonectidae. It belongs to the monotypic genus Nectocaecilia. This species is found in Venezuela and Brazil, and possibly Colombia and Peru. Its natural habitats are subtropical or tropical moist lowland forests and rivers, where it tends to live by the banks of streams. It lives in eel burrows and the roots of floating vegetation. It is threatened by habitat loss. This species has a number of distinct traits that illustrate that it is a burrowing vertebrate, including its very small eyes and long, thin body. (Full article...)