2025 in arthropod paleontology

In 2025, several new arthropod fossil taxa, including arachnids, crustaceans, trilobites, and other arthropods (for insects, see 2025 in paleoentomology) were announced or described. Other significant arthropod paleontological discoveries and events also occurred in 2025.

Chelicerates

Arachnids

Amblypygi

Name Novelty Status Authors Age Type locality Location Notes Image

Phrynus luisdearmasi[1]

Sp. nov

Valid

Dunlop & Bartel

Miocene (probably Burdigalian)

Dominican amber

Dominican Republic

A species of Phrynus.

Phrynus poinari[1]

Nom. nov

Valid

Dunlop & Bartel

Miocene

Mexican amber

Mexico

A species of Phrynus; a replacement name for Phrynus mexicana Poinar & Brown (2004).

Pseudoscorpiones

Name Novelty Status Authors Age Type locality Location Notes Image

Lechytia finniae[2]

Sp. nov

Hagen et al.

Late Cretaceous (Cenomanian)

Kachin amber

Myanmar

A species of Lechytia.

Sarcoptiformes

Name Novelty Status Authors Age Type locality Location Notes Image

Histiogaster altilis[3]

Sp. nov

Valid

Kolesnikov et al.

Eocene

Rovno amber

Ukraine

A species of Histiogaster.

Paralycus ekaterinae[4]

Sp. nov

Valid

Kolesnikov et al.

Eocene (Priabonian)

Rovno amber

Ukraine

A member of Oribatida belonging to the family Pediculochelidae.

Paralycus primus[4]

Sp. nov

Valid

Kolesnikov et al.

Late Cretaceous (Cenomanian)

Kachin amber

Myanmar

A member of Oribatida belonging to the family Pediculochelidae.

Plesioglyphus[5]

Gen. et sp. nov

Valid

Sendi et al.

Early Cretaceous (Barremian)

Lebanese amber

Lebanon

A member of the family Schizoglyphidae. The type species is P. lebanotermi.

Sarcoptiform research
  • Klimov et al. (2025) revise Protospeleorchestes pseudoprotacarus, Paraprotacarus hirsti and Palaeotydeus devonicus from the Devonian Rhynie chert (United Kingdom) and interpret them all as junior synonyms of Protacarus crani, assigned by the authors to the new family Protoacaridae within Endeostigmata; the authors also study the diversification timeline of acariform mites, and argue that the crown group of Acariformes originated during the Cambrian, at the time of colonization of lands by bryophytes.[6]

Scorpiones

Name Novelty Status Authors Age Type locality Location Notes Image

Archaeoananteroides carusoi[7]

Sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Kachin amber

Myanmar

A scorpion belonging to the superfamily Buthoidea and the family Ananteridae.

Burmesescorpiops wunpawng[8]

Sp. nov

Valid

Lourenço, Dan & Zawgyi

Cretaceous

Kachin amber

Myanmar

A scorpion belonging to the family Palaeoeuscorpiidae.

Cretaceoushormiops petersi[9]

Sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Kachin amber

Myanmar

A scorpion belonging to the family Protoischnuridae.

Jeholia[10]

Gen. et sp. nov

Valid

Xuan et al.

Early Cretaceous

Yixian Formation

China

A member of Buthida of uncertain affinities. The type species is J. longchengi.

Palaeoburmesebuthus sarahmeyerae[11]

Sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Kachin amber

Myanmar

A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.

Palaeobutheolus[12]

Gen. et sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Kachin amber

Myanmar

A scorpion, possibly a member of the family Buthidae. The type species is P. andreschmidti.

Paratrilineatus[13]

Gen. et sp. nov

Valid

Lourenço in Lourenço & Velten

Cretaceous

Kachin amber

Myanmar

A scorpion belonging to the superfamily Buthoidea and the family Palaeotrilineatidae. The type species is P. schmidti.

Serratochaerilobuthus barbarae[14]

Sp. nov

Valid

Lourenço & Velten

Cretaceous

Kachin amber

Myanmar

Scorpion research
  • Xuan et al. (2025) revise scorpions from the family Chaerilobuthidae known from the Cretaceous Kachin amber from Myanmar, reinterpret Chaeriloiurus and Serratochaerilobuthus as junior synonyms of the genus Chaerilobuthus, and rerank the family Chaerilobuthidae itself as a subfamily belonging to the family Pseudochactidae.[15]

Trombidiformes

Name Novelty Status Authors Age Type locality Country Notes Images

Caeculus aeternus[16]

Sp. nov

Porta et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Caeculidae.

Cretachyzeria[17]

Gen. et sp. nov

Valid

Liu, Fan & Ren

Late Cretaceous (Cenomanian)

Kachin amber

Myanmar

A member of the family Chyzeriidae. The type species is C. macroseta.

Trombidiform research
  • New caeculid specimens, including the first fossil caeculid larva, are described from the Cretaceous Kachin amber from Myanmar by Gerbe et al. (2025).[18]

Uropygi

Name Novelty Status Authors Age Type locality Country Notes Images

Mesothelyphonus xiaoae[19]

Sp. nov

Wu et al.

Cretaceous

Kachin amber

Myanmar

A whip scorpion.

Eurypterids

Name Novelty Status Authors Age Type locality Country Notes Images

Athenepterus[20]

Gen. et comb. nov

Valid

Lamsdell

Silurian (Přídolí)

United Kingdom

A member of the family Hardieopteridae. The type species is "Eurypterus" megalops Salter (1859); genus also includes A. sigmoidalis (Kjellesvig-Waering, 1971).

Barusopterus[20]

Gen. et comb. nov

Valid

Lamsdell

Silurian (Ludlow)

United States
( Indiana)

A member of Dolichopteroidea belonging to the family Strobilopteridae. The type species is "Onychopterus" limuloides Kjellesvig-Waering (1948).

Cruinnopterus[20]

Gen. et comb. nov

Valid

Lamsdell

Silurian (LlandoveryWenlock)

Canada
( Ontario)
United Kingdom

A member of the family Carcinosomatidae. The type species is "Eurypterus" scorpioides Woodward (1868); genus also includes C. scotica (Laurie, 1899).

Hunanopterus[20]

Fam. et gen. et comb. nov

Valid

Lamsdell

Silurian (Llandovery–Přídolí)

China

A member of the superfamily Pterygotoidea, the type genus of the new family Hunanopteridae. The type species is "Hughmilleria" wangi Tetlie, Selden & Ren (2007).

Selkiepterella[20]

Gen. et comb. nov

Valid

Lamsdell

Silurian (Llandovery–Přídolí)

United Kingdom

A member of the family Adelophthalmidae. The type species is "Himantopterus" lanceolata Salter (1856); genus also includes S. cephalaspis (Salter in Sedgwick and McCoy, 1855).

Tigrisopterus[21]

Gen. et sp. nov

Wang, Sun & Zhang

Devonian (Lochkovian)

Nagaoling Formation

China

A member of the family Carcinosomatidae. The type species is T. zengi.

Waterstonopterus[20]

Gen. et comb. nov

Valid

Lamsdell

Silurian (Llandovery–Wenlock)

United Kingdom
United States
( Pennsylvania)

A member of the family Hardieopteridae. The type species is "Hardieopterus" lanarkensis Waterston (1979); genus also includes W. myops (Clarke, 1907).

Xiphosurans

Name Novelty Status Authors Age Type locality Country Notes Images

Ciurcalimulus[22]

Gen. et sp. nov

Valid

Lamsdell

Silurian (Ludlow)

Wabash Formation

United States
( Indiana)

The type species is C. discobolus.

Lunataspis gundersoni[23]

Sp. nov

Lamsdell et al.

Ordovician (Katian)

Big Hill Formation

United States
( Michigan)

Bears an unusually elongated shovel-shaped prosoma with recurved spines.

Xiphosuran research

  • Evidence indicating that fusion of the opisthosomal segments happened once in the common ancestor to Xiphosura, while loss of visible segment boundaries happened several times during the xiphosuran evolution, is presented by Lamsdell & Ocon (2025).[24]

Other chelicerates

Other chelicerate research

  • Strausfeld, Andrew & Hirth (2025) report evidence of organization of prosoma and cerebrum of Mollisonia symmetrica similar to those observed in extant arachnids, and intepret Mollisonia as likely to be a stem-arachnid phylogenetically close to the arachnid crown group.[25]
  • Lustri et al. (2025) propose that chelicerates belonging to the family Offacolidae were suspension feeders.[26]

Crustaceans

Malacostracans

Name Novelty Status Authors Age Type locality Country Notes Images

Alpheus indicus[27]

Sp. nov

Valid

Amal, Kapur & Prasanna

Miocene (Burdigalian)

Quilon Formation

India

A species of Alpheus.

Arcticocarcinus americanus[28]

Sp. nov

Valid

Schweitzer

Eocene

Vincentown Formation

United States
( New Jersey)

A crab belonging to the family Necrocarcinidae.

Aridelocaris[29]

Gen. et sp. nov

Schweitzer & Schram

Carboniferous (Tournaisian)

United States
( Ohio)

A member of Phyllocarida belonging to the family Sairocarididae. The type species is A. ohioensis.

Cretatrizocheles rodfeldmanni[30]

Sp. nov

Valid

Gašparič et al.

Early Cretaceous (Aptian or Albian)

Slovenia

A hermit crab belonging to the family Pylochelidae.

Distefania pimientai[31]

Comb. nov

Valid

(Ferratges & Zamora in García-Penas et al.)

Early Cretaceous (Aptian)

Spain

A member of the family Dromiidae belonging to the subfamily Goniodromitinae; moved from Palaeodromites pimientai Ferratges & Zamora in García-Penas et al. (2023).

Dysopodus[32]

Gen. et sp. nov

Valid

Schädel et al.

Early Cretaceous (Barremian)

Lebanon

An isopod belonging to the group Cymothoida. The type species is D. gezei.

Eomunidopsis rigolleti[31]

Sp. nov

Valid

Ossó et al.

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of the family Galatheidae.

Feldmannicarcinus[28]

Gen. et comb. et sp. nov

Valid

Schweitzer

Eocene

Castle Hayne Formation

United States
( New Jersey
North Carolina)

A crab belonging to the family Tumidocarcinidae. The type species is "Glyphithyreus" sturgeoni Feldmann et al. (1998); genus also includes new species F. hajzeri.

Grimothea nishioi[33]

Comb. nov

Valid

(Karasawa)

Miocene

Japan

A species of Grimothea; moved from Munida nishioi Karasawa (1993).

Grimothea ogaensis[33]

Comb. nov

Valid

(Hatai & Kotaka)

Miocene

Japan

A species of Grimothea; moved from Kazuoia ogaensis Hatai & Kotaka (1970).

Hyphalocarcinus[34]

Gen. et comb. nov

Valid

Schweitzer, Klompmaker & Luque

Paleocene

Mexico
United States
( Alabama
Texas)

A crab belonging to the superfamily Carpilioidea and the family Palaeoxanthopsidae. The type species is "Dromilites" americana Rathbun (1935).

Karumballichirus khadroensis[35]

Comb. nov

Valid

(Hyžný & Charbonnier in Hyžný et al.)

Paleocene (Danian)

Khadro Formation

Pakistan

A member of Axiidea belonging to the family Callichiridae; moved from Neocallichirus khadroensis Hyžný & Charbonnier in Hyžný et al. (2016).

Karumballichirus lakhraensis[35]

Comb. nov

Valid

(Hyžný & Charbonnier in Hyžný et al.)

Paleocene (Thanetian)?–Eocene (Ypresian)

Lakhra Formation

Pakistan

A member of Axiidea belonging to the family Callichiridae; moved from Neocallichirus lakhraensis Hyžný & Charbonnier in Hyžný et al. (2016).

Karumballichirus maximus[35]

Comb. nov

Valid

(Milne-Edwards)

Holocene

Thailand

A member of Axiidea belonging to the family Callichiridae; moved from Callianassa maxima Milne-Edwards (1870).

Lobetelson feldmanni[36]

Sp. nov

Valid

Klompmaker et al.

Carboniferous (Kasimovian)

Atrasado Formation

United States
( New Mexico)

A member of Belotelsonidea.

Macrocheira enoptra[37]

Sp. nov

Valid

Schweitzer

Eocene

Lookingglass Formation

United States
( Oregon)

A species of Macrocheira.

Mioapseudes[38]

Gen. et sp. nov

Valid

Wallaard

Miocene

Cyprus

A tanaidacean. The type species is M. mediterraneus.

Morellosia[31]

Gen. et sp. nov

Valid

Ossó et al.

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of the family Etyiidae. The type species is M. maestratensis.

Nahecaris carlsi[39]

Sp. nov

Zamora et al.

Devonian (Pragian)

Santa Cruz Formation

Spain

A member of Phyllocarida belonging to the family Rhinocarididae.

Oncopareia californiana[40]

Sp. nov

Valid

Garassino, Pasini & Nyborg

Late Cretaceous (Turonian)

Ladd Formation

United States
( California)

A lobster.

Pelorophontes[41]

Gen. et sp. nov

Schweitzer & Schram

Carboniferous (Moscovian)

United States
( Oklahoma)

A member of the family Gorgonophontidae. The type species is P. mayi.

Pseudoplakolana[32]

Gen. et comb. nov

Valid

Schädel et al.

Early Cretaceous (Aptian)

Sierra Madre Formation

Mexico

An isopod belonging to the group Cymothoida. The type species is "Plakolana" chiapaneca Bruce et al. (2021).

Rodneyellus[42]

Gen. et sp. nov

Valid

Lima et al.

Pirabas Formation

Brazil

A crab belonging to the family Hexapodidae. Genus includes new species R. feldmanni.

Sabellidromites santamarta[43]

Sp. nov

Valid

Lima et al.

Late Cretaceous (Santonian–Campanian)

Santa Marta Formation

Antarctica

A crab belonging to the family Dynomenidae.

Serraranina[44]

Gen. et sp. nov

Valid

Van Bakel, Fraaije & Jagt

Paleocene (Danian)

Denmark
Germany

A raninid crab. Genus includes new species S. stefanpolkowskyi.

Synoriacarcinus[45]

Gen. et comb. nov

Valid

Schweitzer, Feldmann & Findling

Late Cretaceous (Turonian to Campanian)

Merchantville Formation

United States
( Delaware)

A crab belonging to the family Ibericancridae. The type species is "Diaulax" millerae Bishop (1992); genus also includes "Seorsus" kauffmani Feldmann et al. (2013) from the Mancos Shale (New Mexico, United States).

Trapezionida chiyoensis[33]

Sp. nov

Valid

Karasawa & Kato

Miocene

Japan

A species of Trapezionida.

Viapagurus covavidrensis[31]

Sp. nov

Valid

Ossó et al.

Early Cretaceous (Aptian)

Forcall Formation

Spain

A member of the family Paguridae.

Malacostracan research

  • Bicknell et al. (2025) study two clusters of Archaeoniscus brodiei from the Berriasian Durlston Formation (United Kingdom), providing new information on the anatomy of the studied isopod.[46]
  • New information on the morphology of Beurlenia araripensis is provided by Lima et al. (2025).[47]
  • Mychko (2025) describes fossil material of Palaeastacus aff. solitarius from the Tithonian strata from the Cheryomukha River Basin (Yaroslavl Oblast, Russia), extending known geographical range of Late Jurassic members of the genus Palaeastacus.[48]
  • A study on the distribution and diversity of members of the family Glypheidae throughout their evolutionary history is published by Damborenea et al. (2025).[49]
  • Worthy et al. (2025) identify fossil material (molar ridges of the mandible) of at least three taxa of parastacids from the Miocene Bannockburn Formation (New Zealand), providing evidence of greater diversity of parastacids in New Zealand in the Miocene compared to the present.[50]
  • Baucon et al. (2025) report the discovery of vertical burrows from a new Carnian site from the Travenanzes Formation (Italy), possibly representing the oldest fossil evidence of true crabs reported to date.[51]
  • Mychko, Schweitzer & Feldmann (2025) describe fossil material of Gastrosacus wetzleri and Goniodromites aliquantulus from Oxfordian reef limestones of the North Caucasus (Russia), expanding known geographical range of both taxa.[52]

Ostracods

Name Novelty Status Authors Age Type locality Country Notes Images

Aparchitellina lenis[53]

Sp. nov

Valid

Sobolev

Devonian

Yba Formation

Russia

Aparchitellina reticulata[53]

Sp. nov

Valid

Sobolev

Devonian

Yba Formation

Russia

Aurila costabausaensis[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Aurila daphnidis[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Aurila mazzarinoensis[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Aurila sanctiandreae[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Bairdia dorsoundulata[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Podocopida belonging to the family Bairdiidae.

Bairdia timorleste[56]

Sp. nov

Forel in Forel, McCartain & Haig

Triassic

Timor-Leste

Baltonotella erugospinosa[57]

Sp. nov

Valid

Mcgairy et al.

Ordovician (Katian)

Phu Ngu Formation

Vietnam

A member of Podocopa belonging to the order Beyrichiocopida and the family Aparchitididae.

Bizonidea[58]

Gen. et comb. nov

Valid

Franz, Tesakova & Schweigert

Jurassic

France
Germany

Genus includes "Progonocythere" gublerae Bizon (1958).

Chimaerabinoda[57]

Gen. et sp. nov

Valid

Mcgairy et al.

Ordovician (Katian)

Phu Ngu Formation

Vietnam

A member of Podocopa belonging to the order Beyrichiocopida and the family Aechminidae. The type species is C. reticulofimbriata.

Collibolbina corrugata[57]

Sp. nov

Valid

Mcgairy et al.

Ordovician (Katian)

Phu Ngu Formation

Vietnam

A member of Podocopa belonging to the order Beyrichiocopida and the family Tetradellidae.

Cryptobairdia helenentalia[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Podocopida belonging to the family Bairdiidae.

Cyclocypris bamba[59]

Sp. nov

Díez-Somolinos et al.

Early Cretaceous (Barremian)

Spain

A species of Cyclocypris.

Cypridea marihoni[59]

Sp. nov

Díez-Somolinos et al.

Early Cretaceous (Barremian)

Spain

A member of Cypridoidea belonging to the family Cyprideidae.

Cypridea vatra[59]

Sp. nov

Díez-Somolinos et al.

Early Cretaceous (Barremian)

Spain

A member of Cypridoidea belonging to the family Cyprideidae.

Cytheretta buccheriensis[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Glyptocythere pseudotuberosa[60]

Nom. nov

Valid

Tesakova

Middle Jurassic (Bathonian)

Germany

A replacement name for Glyptocythere tuberosa Brand & Malz (1962).

Hiatobairdia peggy[61]

Sp. nov

Forel

Late Triassic (Carnian)

San Cassiano Formation

Italy

Hungarella aitutu[56]

Sp. nov

Forel in Forel, McCartain & Haig

Triassic

Timor-Leste

Judahella leii[62]

Sp. nov

Forel et al.

Middle Triassic

Guanling Formation

China

Laterophores arrisoris[57]

Sp. nov

Valid

Mcgairy et al.

Ordovician (Katian)

Phu Ngu Formation

Vietnam

A member of Podocopa belonging to the order Beyrichiocopida and the family Bolliidae.

Laterophores reticulatus[57]

Sp. nov

Valid

Mcgairy et al.

Ordovician (Katian)

Phu Ngu Formation

Vietnam

A member of Podocopa belonging to the order Beyrichiocopida and the family Bolliidae.

Lophocythere? jhuranensis[63]

Sp. nov

Valid

Kumari & Mahalakshmi in Mahalakshmi, Kumari & Muduli

Late Jurassic (Kimmeridgian)

Jhuran Formation

India

Published online in 2025, but the issue date is listed as December 2024.

Lophocythere mosaica[64]

Sp. nov

Valid

Shurupova

Middle Jurassic (Callovian)

Russia
( Kursk Oblast)

Lophocythere tuberculata[64]

Sp. nov

Valid

Shurupova

Middle Jurassic (Callovian)

Russia
( Kursk Oblast
Ryazan Oblast)

Macrodentina jaduraensis[63]

Sp. nov

Valid

Kumari & Mahalakshmi in Mahalakshmi, Kumari & Muduli

Late Jurassic (Kimmeridgian)

Jhuran Formation

India

Published online in 2025, but the issue date is listed as December 2024.

Mongolocypris shai[65]

Sp. nov

Valid

Wang

Early Cretaceous

Xiaonangou Formation

China

Neurocythere margaritae[66]

Sp. nov

Valid

Tesakova

Middle Jurassic (Callovian)

Russia

Neurocythere parva[66]

Sp. nov

Valid

Tesakova

Middle Jurassic (Callovian)

Russia

Ordovizona? constricta[57]

Sp. nov

Valid

Mcgairy et al.

Ordovician (Katian)

Phu Ngu Formation

Vietnam

A member of Podocopa belonging to the order Beyrichiocopida and the family Ordovizonidae.

Patellacythere lineata[63]

Sp. nov

Valid

Kumari & Mahalakshmi in Mahalakshmi, Kumari & Muduli

Late Jurassic (Kimmeridgian)

Jhuran Formation

India

Published online in 2025, but the issue date is listed as December 2024.

Pattersoncypris labiata[67]

Sp. nov

Maia et al.

Cretaceous

A member of the family Cyprididae.

Perissocytheridea (Kroemmelbeinella) hiblaea[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Polycope latitudialis[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Halocyprida belonging to the family Polycopidae.

Polycope marginoplanata[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Halocyprida belonging to the family Polycopidae.

Polycope marginotorata[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Halocyprida belonging to the family Polycopidae.

Pontopolycope reiflingia[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Halocyprida belonging to the family Polycopidae.

Reconcavona grandiensis[68]

Comb. nov

Valid

(Tomé, Lima Filho & Neumann)

Early Cretaceous

Araripe Basin

Brazil

A member of the family Paracyprididae; moved from Damonella grandiensis Tomé, Lima Filho & Neumann (2014).

Tenedocythere forticostata[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Terquemula pseudoflexicosta[66]

Sp. nov

Valid

Tesakova

Middle Jurassic (Callovian)

Russia

Triadohealdia cribasensis[56]

Sp. nov

Forel in Forel, McCartain & Haig

Triassic

Timor-Leste

Triadohealdia manatuto[56]

Sp. nov

Forel in Forel, McCartain & Haig

Triassic

Timor-Leste

Urobairdia umbonata[55]

Sp. nov

Mette & Moser

Middle Triassic (Ladinian)

Reifling Formation

Austria

A member of Podocopida belonging to the family Bairdiidae.

Verrucocythereis verrucomurata[54]

Sp. nov

Sciuto, Baldanza & Reitano

Pliocene

Italy

Ostracod research

  • Evidence from the study of Silurian ostracod assemblages from the eastern Baltic Basin (Lithuania), indicating that the mid-Homerian biotic turnover event most likely lasted approximately 260,000 years (and thus was shorter than indicated by earlier estimates), is presented by Rinkevičiūtė et al. (2025).[69]
  • Wang et al. (2025) revise the ostracod fauna from the Upper Cretaceous Liwaxia and Madongshan formations (China), correlate it with contemporaneous ostracod faunas from China and Mongolia, and assign the genus Liupanshania to the subfamily Cyproidinae in the family Notodromadidae.[70]
  • A study on the composition and biogeographical connections of ostracod assemblages from the Paleocene-Eocene sedimentary succession at Wadi Tarfa (Egypt) is published by Samir et al. (2025).[71]

Thecostracans

Name Novelty Status Authors Age Type locality Country Notes Images

Alabamalepas[72]

Gen. et sp. nov

Valid

Perreault, Collareta & Buckeridge

Oligocene

United States
( Alabama
South Carolina)

A barnacle belonging to the family Platylepadidae. Genus includes new species A. cookei.

Amphibalanus itoigawai[73]

Sp. nov

Valid

Karasawa & Kobayashi

Early-Middle Miocene (Burdigalian)-(Langhian)

Toyama Formation, Iwamura Groupp; Akeyo and Oidawara formations, Mizunami Group

Japan

A member of the family Balanidae.

Bognorscalpellum[74]

Gen. et comb. nov

Valid

Gale

Eocene

United Kingdom

A barnacle. The type species is "Mitella" venablesi Withers (1953); genus also includes "Mitella" daviesi Withers (1953).

Lacrymascalpellum[74]

Gen. et comb. nov

Valid

Gale

Miocene to Pleistocene

Austria
Czech Republic
France
Italy
 Switzerland
United Kingdom

A barnacle. The type species is "Scalpellum" burdigalense Des Moulins (1875); genus also includes "Scalpellum" magnum Darwin (1851), "Scalpellum" dalpiazi Withers (1953), "Scalpellum" molinianum Seguenza (1876), "Scalpellum" lovisatoi De Alessandri (1895), "Scalpellum" studeri Tièche (1905), "Scalpellum" moraviense Withers (1953) and "Scalpellum" sigmoideum Withers (1953).

Laeviscalpellum[74]

Gen. et comb. nov

Valid

Gale

Cretaceous

United Kingdom
United States

A barnacle. The type species is "Virgiscalpellum" laevis Gale (2020); genus also includes "Virgiscalpellum" rugosum Gale (2020).

Lepas tanakai[73]

Sp. nov

Valid

Karasawa & Kobayashi

Early Miocene (Burdigalian)

Akeyo Formation, Mizunami Group

Japan

A member of the family Lepadidae.

Megabalanus yamaguchii[73]

Sp. nov

Valid

Karasawa & Kobayashi

Early-Middle Miocene (Burdigalian)- (Langhian)

Akeyo and Oidawara formations, Mizunami Group

Japan

A member of the family Balanidae.

Membranobalanus distortus[75]

Sp. nov

Valid

Gale & Sadorf

Pliocene (Piacenzian)

Yorktown Formation

United States
( North Carolina)

A member of the family Balanidae.

Proverruca cenomanica[74]

Sp. nov

Valid

Gale

Late Cretaceous

United Kingdom

A barnacle.

Proverruca minuta[74]

Sp. nov

Valid

Gale

United Kingdom

A barnacle.

Proverruca ornata[76]

Sp. nov

Valid

Gale & Jagt

Late Cretaceous (Campanian)

Gulpen Formation

Belgium

A barnacle.

Solidobalanus hataii[73]

Sp. nov

Valid

Karasawa & Kobayashi

Early-Middle Miocene (Burdigalian)-(Langhian)

Akeyo and Oidawara formations, Mizunami Group

Japan

A member of the family Balanidae.

Solidoscalpellum[74]

Gen. et comb. nov

Valid

Gale

Cretaceous

Kazakhstan
United Kingdom

A scalpellid barnacle. The type species is "Scalpellum maximum" var. cylindraceum Darwin (1851); genus also includes "Arcoscalpellum maximum" var. triminghamensis Withers (1935) and "Cretiscalpellum" sharapovi Alekseev (2009).

Striascalpellum harnhamensis[74]

Sp. nov

Valid

Gale

United Kingdom

A barnacle.

Tetraclitella mizunamiensis[73]

Sp. nov

Valid

Karasawa & Kobayashi

Early Miocene (Burdigalian)

Akeyo Formation, Mizunami Group

Japan

A member of the family Tetraclitidae.

Tetraclitella tokiensis[73]

Sp. nov

Valid

Karasawa & Kobayashi

Early Miocene (Burdigalian)

Akeyo Formation, Mizunami Group

Japan

A member of the family Tetraclitidae.

Tetrinis schooni[77]

Sp. nov

Buckeridge & Smith

Early Cretaceous (Albian)

Allaru Formation

Australia

A barnacle belonging to the group Scalpellomorpha and the family Zeugmatolepadidae.

Titanolepas curvatus[74]

Sp. nov

Valid

Gale

United Kingdom

A barnacle.

Virgilepas angulosum[74]

Sp. nov

Valid

Gale

United Kingdom

A barnacle.

Virgiscalpellum brydonei[74]

Sp. nov

Valid

Gale

United Kingdom

A barnacle.

Thecostracan research

Other crustaceans

Name Novelty Status Authors Age Type locality Country Notes Images

Missouriclus[79]

Gen. et comb. nov

Schweitzer, Mychko & Lehnert

Carboniferous (Kasimovian)

Iola Formation

United States
( Missouri)

A member of Cyclida. Genus includes "Cyclus" communis Rogers (1902).

Ohioclus[79]

Gen. et comb. nov

Schweitzer, Mychko & Lehnert

Carboniferous (Kasimovian)

United States
( Ohio)

A member of Cyclida. Genus includes O. columbianensis.

Oonocarcinus uralicus[80]

Sp. nov

Valid

Mychko, Alekseev & Schweitzer

Carboniferous (Gzhelian)–Permian (Asselian)

Russia
( Perm Krai)

A member of Cyclida.

Palaeolimnadia stevenbeckeri[81]

Sp. nov

Valid

Poschmann et al.

Devonian (Emsian)

Klerf Formation

Luxembourg

A clam shrimp belonging to the family Paleolimnadiidae.

Pavelskine[79]

Gen. et comb. nov

Schweitzer, Mychko & Lehnert

Carboniferous (Kasimovian)

Iola Formation

United States
( Missouri)

A member of Cyclida. Genus includes "Cyclus" packardi Rogers (1902).

Polygrapta dazuensis[82]

Comb. nov

(Chen)

Late Triassic

Xujiahe Formation

China

A clam shrimp belonging to the family Polygraptidae; moved from Euestheria dazuensis Chen (1974)

Polygrapta yipinglangensis[82]

Comb. nov

Valid

(Chen)

Late Triassic

Ganghaizi Formation

China

A clam shrimp belonging to the family Polygraptidae; moved from Euestheria yipinglangensis Chen (1974)

Uralocyclus feldmanni[80]

Sp. nov

Valid

Mychko, Alekseev & Schweitzer

Carboniferous (Viséan or Serpukhovian)

Russia
( Chelyabinsk Oblast)

A member of Cyclida.

Other crustacean research

Insects

Radiodonts

Name Novelty Status Authors Age Type locality Country Notes Images

Mosura[84]

Gen. et sp. nov

Valid

Moysiuk and Caron

Cambrian (Wuliuan)

Burgess Shale

Canada
( British Columbia)

A member of the family Hurdiidae, likely sister to all other genera within the clade. The type species is M. fentoni.

Radiodont research

  • Evidence from the study of new fossil material of Caryosyntrips from the Cambrian strata of the Hongjiangshao Formation (China) and Spence Shale (Utah, United States), interpreted as indicating that characters used to diagnose species belonging to this genus might instead reflect variation within the same species, is presented by Yang et al. (2025).[85]
  • Luo et al. (2025) describe fossil material of Ursulinacaris cf. U. grallae from the Miaolingian strata of the Kaili Formation (China), expanding known geographical range of members of the genus Ursulinacaris.[86]

Trilobites

Name Novelty Status Authors Age Type locality Country Notes Images

Amphoton bicknelli[87]

Sp. nov

Valid

Smith et al.

Cambrian (Miaolingian)

Tasman Formation

New Zealand

A member of the family Dolichometopidae.

Atops antiatlasensis[88]

Sp. nov

Geyer & Landing

Cambrian

Morocco

"Balticoglaucus" avalonensis[89]

Sp. nov

Valid

Westrop & Landing

Cambrian (Drumian)

Manuels River Formation

Canada
( New Brunswick)

A member of the family Solenopleuridae.

Batocara tuberosus[90]

Sp. nov

Valid

Wei et al.

Ordovician-Silurian transition

China

Bergeronites leishuae[91]

Sp. nov

Peng et al.

Cambrian (Guzhangian)

Longha Formation

China

A member of the family Damesellidae.

Catillicephalites? ambiguus[92]

Sp. nov

Valid

Westrop & Dengler

Cambrian

Canada

Ciliscutellum rhaxerosides[90]

Sp. nov

Valid

Wei et al.

Ordovician-Silurian transition

China

Cummingella europa[93]

Sp. nov

Valid

Müller

Carboniferous (Viséan)

Heiligenhaus Formation

Germany

A member of the family Phillipsiidae.

Cummingella pauli[93]

Sp. nov

Valid

Müller

Carboniferous (Viséan)

Heiligenhaus Formation

Germany

A member of the family Phillipsiidae.

Cyrtosymboloides koenigshofi[94]

Sp. nov

Valid

Flick

Devonian

Hermershausen Limestone

Germany

Dinea[92]

Gen. et sp. nov

Valid

Westrop & Dengler

Cambrian (Drumian)

Canada

A probable member of the family Asaphiscidae. Genus includes new species D. bulla.

Ibexocephala[95]

Gen. et 2 sp. nov

Valid

Adrain

Ordovician (Tremadocian)

Garden City Formation

United States
( Idaho)

A member of the family Bathyuridae. The type species is I. lossoae; genus also includes I. dekosterae from the Fillmore Formation (Utah, United States).

Imighzeria[88]

Gen. et sp. nov

Geyer & Landing

Cambrian

Morocco

Genus includes new species I. silena.

"Jincella" arenata[89]

Sp. nov

Valid

Westrop & Landing

Cambrian (Drumian)

Manuels River Formation

Canada
( New Brunswick)

A member of the family Solenopleuridae.

Kayseraspis rugosa[96]

Sp. nov

Valid

Ghobadi Pour

Ordovician (Tremadocian)

Iran

Kettneraspis dickinsoni[97]

Sp. nov

Valid

Van Viersen, Lerouge & Kesselaer

Devonian (Pragian)

Ihandar Formation

Morocco

A member of the family Odontopleuridae.

Leonaspis burketti[97]

Sp. nov

Valid

Van Viersen, Lerouge & Kesselaer

Devonian (Pragian)

Ihandar Formation

Morocco

A member of the family Odontopleuridae.

Leonaspis chancellori[97]

Sp. nov

Valid

Van Viersen, Lerouge & Kesselaer

Devonian (Pragian)

Ihandar Formation

Morocco

A member of the family Odontopleuridae.

Licnocephala bradleyi[95]

Sp. nov

Valid

Adrain

Ordovician (Tremadocian)

Garden City Formation

United States
( Idaho)

A member of the family Bathyuridae.

Licnocephala ngi[95]

Sp. nov

Valid

Adrain

Ordovician (Tremadocian)

Garden City Formation

United States
( Idaho)

A member of the family Bathyuridae.

Lorrettina diemenensis[98]

Sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Maladioidella spinosa[98]

Sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Mansuyites vivianensis[98]

Sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Meadowtownella yunnanensis[90]

Sp. nov

Valid

Wei et al.

Ordovician-Silurian transition

China

Neoacrocephalina[98]

Gen. et sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Genus includes new species N. banksi.

Neogriffithides artamonovorum[99]

Sp. nov

Valid

Mychko

Permian (Wuchiapingian)

Russia
( Primorsky Krai)

Novoriocephalus[98]

Gen. et sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Genus includes new species N. lagoonensis.

Onchonotellus arealis[100]

Sp. nov

Valid

Makarova et al.

Cambrian

Russia

Onchonotopsis formosa[92]

Sp. nov

Valid

Westrop & Dengler

Cambrian

Canada

Paraphillipsia urushtensis[101]

Sp. nov

Valid

Mychko

Permian (Changhsingian)

Belalabino Group

Russia
( Krasnodar Krai)

A member of the family Phillipsiidae.

Parasolenopleura siberica[100]

Sp. nov

Valid

Makarova et al.

Cambrian

Russia

Piriproetus ornatissimus nictans[94]

Ssp. nov

Valid

Flick

Devonian

Hermershausen Limestone

Germany

Politicurus edwardsi[102]

Sp. nov

Valid

Ng, Bradley & Adrain in Adrain et al.

Ordovician (Tremadocian)

House Limestone

United States
( Utah)

A member of the family Hystricuridae.

Praepatokephalus housensis[102]

Sp. nov

Valid

Adrain & Pérez-Peris in Adrain et al.

Ordovician (Tremadocian)

House Limestone

United States
( Utah)

A member of the family Remopleurididae.

Pseudanomocarina falcata[100]

Sp. nov

Valid

Makarova et al.

Cambrian

Russia

Ptychopleurites myrowi[103]

Sp. nov

Valid

Taylor, Strauss & Repetski

Cambrian

Jones Ridge Limestone

United States
( Alaska)

Sinoencrinurus[90]

Gen. et comb. nov

Valid

Wei et al.

Ordovician-Silurian transition

Lianfeng Formation

China

The type species is "Encrinuroides" zhenxiongensis Sheng (1964); genus also includes "Niuchangella" meitanensis Zhang (1974), "Encrinuroides" yanheensis Yin in Yin & Lee (1978), "Encrinuroides" yichangensis Yi (1978) and "Encrinuroides" yinjiangensis Zhang (1974).

Skullrockicurus[102]

Gen. et sp. nov

Valid

Losso & Adrain in Adrain et al.

Ordovician (Tremadocian)

House Limestone

United States
( Utah)

A member of the family Dimeropygidae. The type species is S. plummeri.

Solenoparia bensoni[87]

Sp. nov

Valid

Smith et al.

Cambrian (Miaolingian)

Tasman Formation

New Zealand

A member of the family Solenopleuridae.

Symphysurina ripperdani[103]

Sp. nov

Valid

Taylor, Strauss & Repetski

Cambrian

Jones Ridge Limestone

United States
( Alaska)

Tasmanosaukia[98]

Gen. et sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Genus includes new species T. pulchra.

Toxotiformis tchopkiensis[100]

Sp. nov

Valid

Makarova et al.

Cambrian

Russia

Vicnepea bentleyi[87]

Sp. nov

Valid

Smith et al.

Cambrian (Miaolingian)

Tasman Formation

New Zealand

A member of the family Nepeidae.

?Wuhuia turuaensis[98]

Sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

Yunnanoproetus[90]

Gen. et sp. nov

Valid

Wei et al.

Ordovician-Silurian transition

China

The type species is Y. shanshuensis.

Trilobite research

  • Evidence of impact of changes in marine redox on the evolution of body size of Cambrian and Ordovician trilobites is presented by Sun et al. (2025).[104]
  • A study on the phylogenetic relationships of cyclopygid trilobites is published by Braddy, Dale & Wang (2025).[105]
  • Revision of the family Olenidae is published by Monti (2025).[106]
  • Crônier, Couette & Laffont (2025) compare the utility of 2D and 3D quantitative analyses for the studies of morphological diversity of phacopid trilobites.[107]
  • Zabini et al. (2025) identify fossil material of Mucronaspis sp. from the Ordovician Iapó Formation, representing the oldest record of a trilobite from Brazil reported to date.[108]
  • A study on the modular organization of the trilobite head, as indicated by data from specimens of Ceraurus pleurexanthemus from the Ordovician (Sandbian) Glens Falls Limestone (New York, United States), is published by Vargas-Parra & Hopkins (2025).[109]
  • Nikolic, Warnock & Hopkins (2025) evaluate the utility of inclusion of both fossils with morphological data and ones only with taxonomic constraints and data on age but without morphological data in the analyses aiming to recover dated phylogeny of extinct taxa, testing their combined approach on trilobites from the group Aulacopleurida, and argue that a combined analysis outperforms analyses that only include taxa scored into a morphological matrix.[110]

Other arthropods

Name Novelty Status Authors Age Type locality Country Notes Images

Ammagnostus minutus[100]

Sp. nov

Valid

Makarova et al.

Cambrian

Russia

A member of Agnostida.

?Cotalagnostus novaezealandiae[87]

Sp. nov

Valid

Smith et al.

Cambrian (Miaolingian)

Tasman Formation

New Zealand

A member of Agnostida belonging to the family Spinagnostidae.

Dabashanella? lunaiformis[111]

Sp. nov

Valid

Peel

Cambrian (Wuliuan)

Henson Gletscher Formation

Greenland

A member of Phosphatocopida belonging to the family Dabashanellidae.

Hypagnostus porterensis[89]

Sp. nov

Valid

Westrop & Landing

Cambrian (Drumian)

Manuels River Formation

Canada
( New Brunswick)

A member of Agnostida.

Keurbos[112]

Gen. et sp. nov Valid Gabbott et al. Ordovician

(Hirnantian)

Soom Shale (Cedarberg Formation) South Africa An enigmatic euarthropod, the type species is K. susanae.

Laeviglyphiulus[113]

Gen. et sp. nov

Valid

Wesener & Rühr

Cretaceous

Kachin amber

Myanmar

A millipede belonging to the family Cambalopsidae. The type species is L. patrickmuelleri.

Liangshanella? qassutit[111]

Sp. nov

Valid

Peel

Cambrian (Wuliuan)

Henson Gletscher Formation

Greenland

A member of Bradoriida belonging to the family Svealutidae.

Protosiphonorhinus[114]

Gen. et sp. nov

Valid

Moritz, Wipfler & Wesener

Cretaceous (Albian-Cenomanian)

Kachin amber

Myanmar

A millipede belonging to the family Siphonorhinidae. The type species is P. patrickmuelleri.

Pseudagnostus powenaensis[98]

Sp. nov

Valid

Laurie, Jago & Bischoff

Cambrian (Furongian)

Australia

A member of Agnostida.

Tardisia[115]

Gen. et sp. nov

Valid

McCoy et al.

Upper Carboniferous (Moscovian)

Mazon Creek fossil beds

United States

( Illinois)

A probable late surviving member of the Vicissicaudata within Artiopoda. The type species is T. broedeae

Tasagnostus simpsoni[87]

Sp. nov

Valid

Smith et al.

Cambrian (Miaolingian)

Tasman Formation

New Zealand

A member of Agnostida belonging to the family Diplagnostidae.

Toledodiscus[116]

Gen. et sp. nov

Collantes & Pereira

Cambrian Stage 4

Soleras Formation

Spain

A member of Agnostida belonging to the family Weymouthiidae. The type species is T. valverdi.

  • Collantes & Pates (2025) revise the holotype of Isoxys carbonelli and confirm placement of this species within the genus Isoxys.[117]
  • Nielsen et al. (2025) study changes of elongated cardinal spines of Isoxys volucris during its ontogeny, and argue that the studied spines had a primary defensive function.[118]
  • Haridy et al. (2025) identify purported early vertebrate Anatolepis as an arthropod, and interpret its purported dentine tubules as sensory structures similar to those present in Cambrian aglaspidids and modern arthropods.[119]
  • A study on the morphology of cephalic appendages of Acanthomeridion serratum, providing evidence of probable adaptations to durophagy, is published by Wu et al. (2025).[120]
  • Evidence of the presence of two pairs of different compound eyes in Pygmaclypeatus daziensis (a pair of stalked, movable eyes and a pair of sessile dorsal eyes) is presented by Schmidt et al. (2025).[121]
  • O'Flynn et al. (2025) describe new fossil material of Kuamaia lata from the Cambrian Chiungchussu Formation (China), providing new information on the frontal appendages and number of head segments in members of this species, and interpret the studied fossils as indicating that raptorial frontal appendages, ancestral for Euarthropoda but lost in Artiopoda, evolved secondarily within the artiopod lineage that included K. lata.[122]
  • Redescription and a study on the affinities of Helmetia expansa is published by Losso, Caron & Ortega-Hernández (2025).[123]
  • Bicknell et al. (2025) describe fossil material of Naraoia cf. bertiensis from the Silurian (Přídolí) Phelps Member of the Fiddlers Green Formation (Bertie Group; New York, United States), expanding known geographical range of the youngest naraoiids.[124]
  • Naimark & Chaika (2025) study the structure of the cuticles of members of Agnostina, reporting evidence of greater similarity to cuticles of chelicerates than those of crustaceans.[125]
  • Liu et al. (2025) present new information on the morphology of Primicaris, interpreted as supporting a stem-group mandibulate affinity for marrellomorphs.[126]
  • Brookfield, Catlos & Garza (2025) argue that the strata of the Stonehaven Group (United Kingdom) preserving fossil material of Pneumodesmus newmani is most likely PřídolíLochkovian in age.[127]
  • Dernov (2025) describes impressions of probable paratergites of Arthropleura sp. from the Carboniferous (Bashkirian) Mospyne Formation (Ukraine), possibly representing fossil material of juvenile specimens, and argues that juvenile and adult arthropleurids might have lived in different habitats.[128]

General research

  • Chipman (2025) proposes a new model for the evolution of arthropod tagmata based on data from extant and fossil arthropods.[129]
  • Naimark & Sizov (2025) study the taphonomy of the Cambrian arthropod fossils from the Kimiltei site (Irkutsk Oblast, Russia) first reported by Naimark, Sizov & Khubanov (2023),[130] and argue that the identification of putative members of Offacolidae and Chasmataspidida from this locality as chelicerates is correct.[131]
  • Probable burrows of arthropods (possibly insect larvae) feeding on plant roots, as well as their fossilized fecal pellets, are described from the Middle-Upper Triassic strata in Somerset (United Kingdom) by Howson, Tucker & Whitaker (2025).[132]
  • Le Cadre et al. (2025) report the discovery of a bristly millipede very similar to extant Polyxenus lagurus and two member of an extinct mite lineage (Glaesacarus rhombeus) in a single piece of the Eocene Baltic amber, providing evidence of shared habitat of the studied arthropods and possible evidence of bradytely in bristly millipedes.[133]

References

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  2. ^ Hagen, M.; Kotthoff, U.; Harms, D.; Loria, S. F. (2025). "New species of Lechytia Balzan, 1892 (Arachnida: Pseudoscorpiones) from Burmese amber highlights 99 million years of morphological stasis". Cretaceous Research. 172 106120. Bibcode:2025CrRes.17206120H. doi:10.1016/j.cretres.2025.106120.
  3. ^ Kolesnikov, V. B.; Vorontsov, D. D.; Perkovsky, E. E.; Klimov, P. B. (2025). "An exceptionally well-preserved Eocene fossil mite, Histiogaster altilis sp. n. (Acari: Astigmata), from tree sap: Evidence of morphological and ecological niche conservatism, with a review of fossil Astigmata". Acarologia. 65 (1): 213–241. doi:10.24349/c35e-8bmj.
  4. ^ a b Kolesnikov, V. B.; Vorontsov, D. D.; Norton, R. A.; Klimov, P. B. (2025). "First fossil evidence of pediculochelid mites: two new species from Middle Cretaceous and Late Eocene amber revealing morphological stasis over at least 99 million years". Acarologia. 65 (1): 67–90. doi:10.24349/uxz4-s4sq.
  5. ^ Sendi, H.; Klimov, P. B.; Kolesnikov, V. B.; Káčerová, J.; Bonino, E.; Azar, D.; Robin, N. (2025). "The oldest continuous association between astigmatid mites and termites preserved in Cretaceous amber reveals the evolutionary significance of phoresy". BMC Ecology and Evolution. 25 (1). 16. Bibcode:2025BMCEE..25...16S. doi:10.1186/s12862-025-02351-5. PMC 11853500. PMID 39994514.
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  7. ^ Lourenço, W. R.; Velten, J. (2025). "A new species for the genus Archaeoananteroides Lourenço, 2016, fossil scorpion from Burmite (Scorpiones: Buthoidea: Ananteridae)". Faunitaxys. 13 (3): 1–6. doi:10.57800/faunitaxys-13(03).
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  9. ^ Lourenço, W. R.; Velten, J. (2025). "One more new species for the genus Cretaceoushormiops Lourenço, 2018 from Cretaceous Burmite (Scorpiones: Protoischnuridae)". Faunitaxys. 13 (2): 1–6. doi:10.57800/faunitaxys-13(02).
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